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All of the towers with the full upgrades

Bloons Tower Defense 4 has all the towers from the previous games, including 8 brand new ones, one of which has been confirmed. Note: All prices are the Medium price.

Towers

Dart Monkey

"Shoots a single dart that pops a single Bloon. Can upgrade to a Spike-O-Pult with a level four upgrade." ~ Game description'

Dart Monkey (NP)

Dart Monkey

The Dart Monkey is the most basic tower in the Bloons Tower Defense series. It has been in every BTD game. Hotkey: D Cost: $200 Rank to Unlock: 1

Upgrades

  • Spike-O-Pult Cost: $300 Gives the monkey a catapult that hurls giant spiked balls.
  • Triple Shot Cost: $1,000 Spike-O-Pult becomes three times as deadly. Rank to Unlock: 19(Total Selling Price: $1,384)

Tack Shooter

TackShooter

Tack Shooter

Shoots a volley of tacks in eight directions. Has short range, but is good for thinning out Bloons at intersections and corners. Hotkey: T Cost: $360 Rank to Unlock: 2

Upgrades

  • Extra Range Cost: $100 Increases the attack radius of the Tack Shooter.
  • Blade Shooter Cost: $280 Shoots razor sharp bladed discs instead of tacks!
  • Ring of Fire Cost: $2,500 Throws out a fiery hot burst of flame that can pop anything. Rank to Unlock: 22(Total Selling price: $2,760)

Boomerang Thrower

BoomerangThrower

Boomerang Thrower

Throws a boomerang that follows a curved path back to the tower. Can pop multiple Bloons at once. Hotkey: R Cost: $400 Rank to Unlock: 3

Upgrades

  • Multi Target Cost: $250 Boomerang will hit up to seven Bloons at once.
  • Sonic Boom Cost: $100 Sonic boomerangs smash through frozen Bloons.
  • Glaive Thrower Cost: $280 Glaives slice through up to twelve Bloons at once!
  • Lightsabre Thrower Cost: $1,850 Lightsabre slices through anything and everything up to seventy Bloons at once! Rank to Unlock: 20(Total Selling Price: $2,304)

Bomb Tower

BombTower

A Bomb Tower

Launches a bomb that explodes on impact. Can destroy many Bloons at once. Hotkey: B Cost: $700 Rank to Unlock: 4

Upgrades

  • Longer Range Cost: $300 Can shoot bombs further than normal.
  • Missile Launcher Cost: $210 Shoots fast missiles instead of bombs that go faster, farther, and pop more. (Note: The Missile Launcher upgrade also grants a slight fire rate increase.)
  • MOAB Mauler Cost: $900 Special missiles cause 10x damage to MOABs and BFBs. Rank to Unlock: 17(Total Selling Price: $1928)

Ice Tower

IceTower

An Ice Tower

Freezes nearby Bloons with every pulse. Frozen Bloons are immune to anything sharp. Hotkey: I (Uppercase 'i') Cost: $380 Rank to Unlock: 6

Upgrades

  • Permafrost Cost: $100 Slows Bloons Down even after they thaw out!
  • Snap Freeze Cost: $400 Freezes so violently that Bloons will pop before freezing!
  • Arctic Wind Cost: $6,000 Extreme cold slows down anything that comes near the Ice Tower. Rank to Unlock: 24(Note: When upgraded to Arctic Wind, Bloons that are in the Ice Tower's attack radius are slowed down 66%.) (Total Selling Price: $5,684)

Mortar Tower

MortarTower

A Mortar Tower.

Targets a specific area of ground over any distance with an explosive attack. Hotkey: M Cost: $825 Rank to Unlock: 7

Upgrades

  • Greater Accuracy Cost: $200 Increases the mortar's accuracy.
  • Rapid Reload Cost: $250 Increases the mortar's firing rate.
  • Bloon Buster Cost: $800 Smashes through two layers of Bloons.
  • Mortar Battery Cost: $2,000 Triple the pain. Rank to Unlock: 23(Total Selling Price: $3260)

Glue Gunner

GlueGunner

A Glue Gunner

Shoots a glob a glue at at Bloon, slowing it down. Ceramic Bloons can be glued but will not be slowed down. Hotkey: G Cost: $300 *At time of writing, repeat of the word "at" is indeed in the Glue Gunner's description. Rank to Unlock: 9

Upgrades

  • Stickier Glue Cost: $120 Makes Glue last much longer.
  • Glue Soak Cost: $200 Glue soaks through all layers of Bloons.
  • Corrosive Glue Cost: $300 Glue eats through Bloons, dissolving them every two seconds.
  • Glue Splatter Cost: $3000 Affects a large area around impact, soaking up to twenty Bloons at once. Rank to Unlock: 25(Total Selling Price: $3,136)

Monkey Beacon

MonkeyBeacon

A Monkey Beacon

Increases the attack range of all stuff within the beacon area. Has several useful upgrades that 'help out nearby towers, including its ultimate ability to call in Super Monkey Storms. Hotkey: N Cost: $1,000 Rank to Unlock: 10

Upgrades

  • Wider Influence Cost: $500 Increses area of effect for all the beacons abilities.
  • Jungle Drums Cost: $1,500 Increases attack speed of all towers withing the Monkey Beacon's radius.
  • Sonar Beacon Cost: $280 Allows targeting of Camo Bloons for towers inside the radius.
  • Monkey Storm Cost: $3500 Calls in a wave of Super Monkeys* that wipes out all Bloons on screen. Unique. Rank to Unlock: 18 (Note: After you purchase the Monkey Storm upgrade, a new upgrade will appear called Call Super Monkey Storm, cost $1000. Every time you buy it, every Bloon excluding MOABs and BFBs will be destroyed. But after each use you must wait approx. 65 seconds to use it again.) (Total Selling Price: $5,424)

Monkey Ace

MonkeyAce

A Monkey Ace

The only unit that moves, the Monkey Ace patrols the skies above the action, regularly strafing the area with powerful piercing darts. Hotkey: A Cost: $900 (Note: The Monkey Ace tower moves in a vertical figure-8 that covers the height of roughly the entire screen.) Rank to Unlock: 11

Upgrades

  • Pineapple Express Cost: $200 Drops highly explosive pineapples.
  • Spy Plane Cost: $350 Allows detection of Camo Bloons for all towers near the plane.
  • Rapidfire Cost: $700 Increases attack rate.
  • Operation: Dart Storm Cost: $3,000 Our darts will block out the sun. Rank to Unlock: 27(Note: The Operation: Dart Storm upgrade makes it so the Monkey Ace tower fires twice as many darts, twice as fast.) (Total Selling Price: $4,120)

Monkey Buccaneer

MonkeyBuccaneer

A Monkey Buccaneer

The only unit that can be placed in water - the Monkey Buccaneer has a long range and shoots a single, heavy dart. Hotkey: U Cost: $600 Rank to Unlock: 12

Upgrades

  • Grape Shot Cost: $300 Adds a volley of deadly sharpened grapes to the attack.
  • Crow's Nest Cost: $150 Allows detection of Camo Bloons for any towers in the radius.
  • Longer Canons Cost: $180 Increases attack range of the ship's canons.
  • Battleship Cost: $2,000 Total nautical domination. Rank to Unlock: 21(Note: The Battleship upgrade doubles the attack speed the of Monkey Buccaneer.) (Total Selling Price: $2,584)

Monkey Apprentice

MonkeyApprentice

A Monkey Apprentice

Trained in the ancient arts of monkey wizardry. Learns new spells with every upgrade. This unit is flexible and will continue randomly using his old spells with his new one. Hotkey: W Cost: $550 Rank to Unlock: 14

Upgrades

  • Intense Magic Cost: $300 Wizard's magic orb becomes more powerful, pop* up to seven Bloons at once. *At time of writing, this gramatical error is indeed in this upgrade's description.)
  • Fireball Cost: $300 Hurls a burning ball of flame that explodes on impact
  • Summon Whirlwind Cost: $2,000 Whirlwind* will blow Bloons away from the exit, but will thaw Bloons and remove glue
  • Tempest Tornado Cost: $8,000 Tempest tornados* pop Bloons as well as push them around. Affects up to 120 Bloons at once. Rank to Unlock: 29*At time of writing, this spelling error is indeed in this upgrade's description. (Total Selling Price: $8,920)

Super Monkey

SuperMonkeyBTD4

A Super Monkey

Super Monkey shoots incredibly fast. Can upgrade to Laser Vision, then Plasma Vision. Ultimate upgrade is ultimately powerful. Find out for yourself... Hotkey: S Cost: $4,000 Rank to Unlock: 15

Upgrades

  • Super Range Cost: $1,000 Super Range greatly increases Super Monkey attack radius.
  • Laser Vision Cost: $3,500 Lasers can pop two Bloons at once, and can pop frozen Bloons.
  • Plasma Vision Cost: $4,000 Plasma shoots twice as fast and vaporizes everything it touches
  • Sun God Cost: $20,000 Legends speak of a being that fears no Bloon. Rank to Unlock: 30(Note: The Sun God upgrade shoots a little bit faster and divides into three beams.) (Total Selling Price: 26,000)

Banana Farm

BananaFarm

A Banana Farm

Generates extra money at the end of every round, but doesn't attack anything. Can upgrade to increase money bonus. Hotkey: F Cost: $1,000 Rank to Unlock: 16

Upgrades

  • More Bananas Cost: $400 Increases money generated to one hundred twenty per round.
  • Banana Plantation Cost: $1,200 Banana Plantation generates two hundred fifty money per round.
  • Banana Republic Cost: $2,500 Banana Republic pulls in a cool five hundred money per round.
  • Banana Research Facility Cost: $10,000 High Tech banana facility earns you a staggering two thousand money per round. Rank to Unlock: 28(Total Selling Price: $12,080)

Dartling Gun

DartlingGun

A Dartling Gun

Shoots Darts rapidly but a little inaccurately to wherever your mouse is. You can decide exactly where this tower will shoot by moving your mouse wherever you want! Hotkey: None yet Cost: $1250 Rank to Unlock: 1

Upgrades

Steady Barrels Cost: $250 Steady Barrels reduces the spread of darts fired from the gun.

Powerful Shots Cost: $600 Darts fly out with greater speed, popping up to 3 bloons at once.

Increased Barrel Spin Cost: $2,500 Makes dartling gun fire much faster.

Laser Cannon Cost: $15,000 Converts to a powerful cannon that shoots rapid blasts able to cause massive bloon damage. Rank to Unlock: 33(Total Selling Price: ? )

Spike Factory

Screen shot 2011-09-10 at 1.40

Generates piles of roadspikes around the track automatically. Leftover spikes dissapear after each round. Hotkey: none yet. Cost: $700 Rank to Unlock: 1

Upgrades

Increased Production Cost: $595. Makes the factory produce spikes much faster.

'Bigger Stacks' Cost: $510. The factory makes piles of spikes instead of 5.

White Hot Spikes Cost: $300. White hot spikes can pop frozen and lead bloons.

Spike Storm Cost: $5950 Covers the whole path with spikes that go away over time. Rank to Unlock: 34

Road Items

Road Spikes

Spike

Road Spikes

Place these road spikes on the track to pop Bloons. Can pop 10 Bloons before wearing out. Road Spikes last only until the end of the round. Hotkey: K Cost: $30. Rank to Unlock: 5

Pineapple

Pineapple

A Pineapple in a Blue Background

Like all healthy food, Pineapples explode shortly after being placed, so don't put any down unless you want to blow up some Bloons. Hotkey: P Cost: $25 Rank to Unlock: 8

Monkey Glue

MonkeyGlue

Monkey Glue on a track

Monkey Glue slows down Bloons. Each glob can pop twenty Bloons before wearing out. Monkey Glue only lasts until the end of the round. Hotkey: E Cost: $10 Rank to Unlock: 13

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